Usually, when we talk about VPN, we talk about a “remote access” solution. A user from an external location (on a local network) connects to the corporate network to access resources like an FTP server. So we have a single user on one side and a network on another side. Site-to-site VPN is a model when we have a network on both sides.
Imagine you have a company with two offices - one in London and the second in Berlin. If a user from Berlin needs to access a resource in London, he can use a remote access VPN and connect to the London office. However, we have more users accessing resources in different locations. In that case, it makes more sense to create a site-to-site VPN between the London office and the Berlin office, so anyone in one office can access resources in the second office as if they were on the same network.
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The mesh is a collection of nodes, a lighthouse, and an admin center. The user device is a node, a server is a node, cloud stack is a node, LAN access box is a node.